Pseudophilautus steineri Steiner’s Shrub Frog
Knuckles Forest Reserve, Matale District, Central Province, Sri LankaAugust 6, 2019
Steiner’s Shrub Frog (Pseudophilautus steineri) Steiner’s Shrub Frog (Pseudophilautus steineri) Steiner’s Shrub Frog (Pseudophilautus steineri)
Careful surveys combined with molecular analysis in the 2000's led to a slew of new Pseudophilautus species described in Sri Lanka (though at the time they were placed in Philautus). Pseudophilautus steineri is one of those species, distinguished from its previously known sister species P. microtympanum by some molecular sequence divergences and minor morphological differences.

The frogs in these photos were all identified as P. steineri by my guide Udaya Chanaka.

Knuckles Forest Reserve, Matale District, Central Province, Sri LankaAugust 8, 2019
Steiner’s Shrub Frog (Pseudophilautus steineri)
The wide assortment of colors and patterns make this species particularly hard to identify, at least for me.
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